From original description (Doczkal and Pape 2009).
Head. In frontal view almost circular, in lateral approximately twice as high as long, with a depression
of the postocular orbit and the posterior part of the eye lateral to the dorsal eye corner. Dorsal and ventral halves of eye of about the same size, its posterior margin almost straight, only faintly curved anteriorly above the vortex, holoptic, eye contiguity approximately half the length of frons, the eye rims actually separated by an extremely narrow stripe of the frontal cuticle, anterior angle of eyes ca.
60º, ommatidia near eye contiguity insignificantly enlarged. Eyes rather sparsely setose, the margins and the eye bridge broadly bare, the setae 0.2 mm long, glistening silvery. Cuticle of head capsule (including back) entirely covered by dense microtrichia, which are mostly white, but brown on some part
of the back (especially near occipital foramen). Face concave, anterior mouth edge protruding, tentorial pit slit-like, facial ridge indistinct, border with antennal sockets straight, with a small weak tubercle below each antenna, ground colour blackish, with a reddish tinge laterally, difficult to observe under the dense layer of microtrichia, sparsely covered by short erect white setae laterally, leaving about the median third bare at about half height of face, and receding dorsally and ventrally, with a narrow gap between the facial and the frontal setosity, anterior corners of mouth edge bare. Subcranial cavity wide, anteriorly notched, lateral corner of mouth edge somewhat receding. Anteclypeus wide, about twice as long medially as wide proximally, widening distally, completely microtrichose, postclypeus sclerotized. Labella of proboscis externally without striking peculiarities (compared with Merodon spp.). Frons covered by a dense fringe of long, thick, subappressed, creamy white setae, anteriorly with erect, thin, brown setae, that are decreasing in length ventrally, ventrally reaching level of the ventral margin of antennal sockets, with a strong, point-like pit at half length, cuticle black, except for a triangular, non-setose area between lunule and frontal pit, which is red. Lunule dark brown, with slightly raised margins laterally, somewhat thickened, and a little produced in the middle, where it bears small, microtrichose spot, otherwise without any vestiture, ventrally reaching level of ventral base of antenna. Antennal sockets approximately 1.5 times as wide as deep, without a median sclerotized stripe (see, Hippa & Ståhls, 2005: fig. 5c). Postocular orbit dorsally only little wider than at its narrowest part, without a postcranial carina, dorsally horizontal, below the depression sloping towards the posterior surface of the head, with which it has an angled border, parasagittal sulci well developed, long, white setae present at the posterior margin between the lateral ends of the depression, but leaving the vertex (i.e. the area between the parasagittal sulci) bare, and at about the ventral quarter of head, dark brown setae mixed with short, light setae are present anteriorly, leaving a narrow stripe (about as wide as an ocellus) along the eye margin bare, some of the short setae more or less spinose (without clear distinction to the slender setae). Vertical triangle long and narrow, with evenly diverging margins, its posterior part from the anterior ocellus to the posterior margin of the head distinctly raised above the level of the eyes, ocellar triangle equilateral, distance between posterior ocelli and eye corners equals the length of the ocellar triangle, posterior ocelli elongated (about twice as long as wide), nearly touching eye margin. Vertical triangle rather densely covered with long, dark brown setae, a few white setae present at the anterior end of the setosity, except for a number of bare areas: at the posterior end, where a stripe as long as the width of an ocellus is bare, narrow stripes along the eye margins from the posterior ocelli to the eye corners, the anterior half of the area between the anterior ocellus and the eye contiguity, and a narrow median stripe from the posterior end of the posterior ocelli towards the posterior margin of the vertical triangle. Postvertex narrow, with almost parallel margins, bare, with a deep pit at its ventral end. Hypostomal bridge with a sharp, transverse arciform crest near the ventral margin of the occipital foramen. The dorsoventral bulge wider than usual in Merodon, gradually narrowing ventrally. Upper two-thirds of tempora setose, postgena and hypostomal bridge almost bare, with sparse, inconspicuous, short setae. The bare spot at the eye margin behind the ill-defined genal suture narrow (i.e. as usual within Eumerini, but in contrast to Platynochaetus spp., where this spot is much enlarged). Antenna blackish, postpedicel dorsally reddish, short, pedicel much shorter than deep medially, about as long as deep laterally, postpedicel about as long as deep. Scape about as long as deep, concave ventrally, with a strong bump on median surface. Pedicel laterally with a nearly rectangular excavation of its apical margin, its median surface nearly flat. Setae on scape and pedicel black to dark reddish brown, scape with two long apicoventral setae, the longer one nearly as long as width of scape, a group of five short setae somewhat irregularly arranged near the dorsal apex medially, and a larger group of short setae in line near the apex laterally, pedicel on medial surface almost entirely setose except for a small, bare, subapical area at the distal margin, setae on lateral surface restricted to dorsal and distal third, with a narrow gap between the dorsal and the apical patches, the dorsoapical setae short (less than 1.5 times maximum width of arista), the apicoventral setae as long as about half depth of pedicel. Postpedicel with a well-demarcated dorsal fossette, with a narrow basal section differentiated (but difficult to see, especially laterally, with two to three sensory pits on median and lateral surface close to distal margin of basal section, one of which is as wide as maximum width of arista. Arista inserted dorsally near base, long, with nearly circular cross-section, gradually narrowing from base to apex, dark, reddish brown, microtrichose throughout, aristomere 1 not visible externally (but present), aristomere 2 shorter than wide.
Thorax. Black to dark reddish brown, but scutellum and mediotergite light reddish brown, scutellar rim yellow, entirely densely covered by microtrichia, except for the posterior half of the scutellum, which is largely bare of microtrichia. Postpronotum, notopleuron, anepisternum on posterodorsal half of both parts, and anterior part of anepimeron (except for the posteroventral corner), including the dorsomedian part, with a dense cover of long, thick (but not feathered), glistening golden setae, similar but less bright pile present on postalar callus and scutellum, remaining parts of scutum with rather dense, uneven, almost erect black setae composed of setae of two lengths, the shorter present on entire surface, the longer missing sublaterally, a few black setae present at the posterior end of the notopleuron, the anterior end of the postalar callus, and anteroventral end of the anepisternal setosity, median half of premetacoxal bridge rather densely black setose, sparse and inconspicuous short silvery or black setae are present on the proepimeron, pleurae otherwise bare, notably the katepisternum without any setae, about three to four irregular rows of marginal setae of supraalar area moderately thickened. Postalar wall with a strong, three-sided pyramidal tubercle. Scutoscutellar suture deep and narrow, without any trace of a prescutellum. Anterodorsal margin of anepisternum, at the border with postpronotum, with a dorsally directed peg. Katatergite smooth, with only a small posterodorsal part bearing slightly elongated microtrichia, which seem to be arranged as oblique dorsoventral stripes (Hippa & Ståhls, 2005: character 50, state 1; requires verification). Katepimeron convex. Metathoracic spiracle directed posteriorly, occupying nearly half the distance from the ventral margin of the katatergite to the base of the hind coxa, membranous cleft of metapleural sclerites posterior to spiracle narrow, the sclerite ventral to the cleft posterodorsally incised. Premetacoxal bridge welldeveloped, its posteromedian apex simple. Posterior basalare short, pleural wing process with a short membranous cleft, subalar sclerite with a moderately inflated anterodorsal apex, plumule small, dark brown, apparently with simple setae (requires verification with SEM). A small deep pit at
the posterodorsal corner of the meron. Posteroventral area of anepimeron flat, not sunk, thus forming a unified, flat surface together with the anterior part of the katatergite. Ventral surface of scutellum concave. Subscutellum small. Anterior part of anepisternum convex, only slightly differentiated from the posterior part, the posterior part with a rudimentary posterodorsal swelling. Katepisternum anteriorly and at the anapleural suture unusually deeply invaginated (possibly an artefact from being dried out from alcohol). Proepimeron strongly protruding ventrally. Proepisternum laterally slightly bulging, its ventral part bare, its dorsal part with sparse short more or less spinose setae. The arms of the antepronotal collar touching dorsally, the cleft between them only slightly extended posteriorly. Probasisternum 2.3 times as wide as long, almost bare. Haltere dark brown, except for the median part of the stalk, which is yellow. Wing. Infuscated brown, entirely covered by unusually dense microtrichia. Costa ending far from apex, with a short stump distal to R4+5, M1 bent towards wing margin, then recurrent and joining R4+5 at an obtuse angle, the recurrent section without external spur, M2 present as a very short spur, R4+5 deeply curved into cell r4+5, with an additional crossvein connecting the curve of R4+5 with M1 at the level of maximum proximity, cross-vein Sc-R present, R1 long and reaching C at about two-thirds the distance from Sc to R2+3. Tegula densely setose, costagium densely setose, except for a narrow bare stripe along its posterior margin, setae all black (including ventral ones), C microtrichose, with two rows of setae, except for the basal part from the costagium to about one-third of the second costal section, where there are several irregular rows ventrally, the ventral setae being elongated (up to about three times costal width), and two to three moderately irregular rows dorsally from apical half of first costal section to first third of second section. Humeral cross-vein slightly shifted apically in relation to the ‘scar’ near the base of the radial stem vein. Cross-vein R-M placed at about two-thirds of cell dm, posterior angle of cell dm angular, with a short veinlet (CuA1). Vena spuria reaching R-M as a pigmented (brown) wing fold, depigmented proximal to the slight kink in the middle. A1+CuA2 in proximal half straight and strong, then weakened and smoothly curved towards wing margin. Lower calypter dark brown, without long setae on its disk, marginal fringe of upper part white, composed of rather dense, apparently simple setae (observed with stereomicroscope at 80×), marginal fringe of lower part light brown and composed of multifurcate trichiae. Legs. All parts black or reddish brown, with ill-defined borders between the colours, setosity dense, black to dark brown and generally rather short, except for moderately long setae posterolaterally on fore and mid femora. Fore femur anterobasally with a patch of very dense, short setae. Apex of fore tibia anterolaterally with dense, irregularly placed setae. Fore coxa short, about as long at its median margin as it is wide apically, microtrichose (except for basal rim), anteriorly with black setae. Outer part of ventral margin of anterior mesocoxite rather smoothly folded (compared with Merodon, where the fold is sharp), posterior mesocoxite flat, microtrichose, bare. Hind femur moderately thickened, at its widest point (except for process) 0.28 times as wide as long, lower side slightly concave, with an anterolateral apical triangular flange, which bears slightly thickened setae at its posterior margin, without posteroventral subapical spinose setae, ventrally with a bare stripe from base to apex, with a spot of dense, short, thick setae anterolaterally near base, almost entirely microtrichose, except for small areas at the subapical process, the bare ventral area posterior to the subapical flange slightly depressed. Hind tibia with a blunt, anteroventral, sparsely setose carina for about proximal two fifths, the scar placed in middle of the tibia, where the tibia is thickened, apex ventrally with a sharp transverse edge across nearly full width, almost entirely setose, except for a spot at the dorsal apex, which is bare and bright orange, the setae entirely black, the posteroapical setae of the dorsal half elongated (about three-quarters apical width of hind tibia), nearly entirely microtrichose, except for the anteroventral carina, the apicoventral fifth, and the bare dorsoapical spot. Hind trochanter with a moderate hump, covered by dense, short setae, distally bare. All tarsi simple. Claws large (one claw of
hind tarsus 0.61 mm long), base dark red.
Abdomen. Antetergite almost fused with tergite 1, bare. Tergite 1 light reddish brown, darkened towards the sides, covered by short, black (semi-) adpressed setae, except for the posterior margin, which is bare, but densely microtrichose, moderately folded, and about as long as the width of the first tarsomere of mid leg. Lateral part of tergite 4 extended medioventrally, meeting sternite 4. Tergites 2–4 densely covered by bright golden fur, except for a small patch of black setae at the anteromedian part of tergite 2 and the ventral extension of tergite 4, each with a pair of oblique impressions, those on tergite 3 curved (as in Eumerus spp.), the cuticula entirely matt, the setae placed on small black tubercles. First abdominal spiracle placed in the pleural membrane next to the metepimeron, pleural membrane dark brown. Sternite 1 entirely sclerotized, its anterior corners broadly fused with the metepimeron, with a pair of large, setose areas, lateral margins and median line broadly bare. Sternites 2 and 3 much wider than long, covered with short, black setae, leaving the median line bare, sternite 4 nearly twice as wide as long laterally, lateral margins thickened, posterior margin excavated, the posterolateral margins folded towards the body (i.e. upwards), covered by black setae, which are
very short (much shorter than on sternite 3), but long at the lateral margins, median line with a fine sharp line. Terminalia enormously enlarged, black setose.
Terminalia. Epandrium dorsally short, the depressed area proximal to the cerci almost reaching the proximal margin of epandrium, laterally with an extensive membranous area, but without a triangular membranous incision (as generally found in the Eumerini). Cerci largely sclerotized, fused laterally with epandrium for about two-thirds their length, the sclerite basally with a narrow, membranous cleft. Subepandrial sclerite about twice as long as wide, its posterior third heavily sclerotized and with a pair of medially directed processes, at the margins, close to the attachment of surstyli, with a pair of smaller processes, the median line membranous as is the most part of the proximal two-thirds, which are strongly sclerotized laterally and at the anterior margins only. Surstylus firmly fused with epandrium, with well-developed suture between them, posterior lobe (i.e. posterior to the attachment to epandrium) small, its apex curved ventrally, and with a sharp transverse edge, except for the posterior end rather densely clothed with short, black, somewhat spine-like setae, the remaining part of surstylus smooth, bare, except for a small area at its posterior end medially, which has an extension of the setosity of the posterior surstyle. Hypandrium with an extraordinarily voluminous base, the anterodorsal corners, which are attached to the anteroventral corners of the epandrium, long and slender, as is the dorsomedian process, with which the hypandrium is attached to the subepandrial sclerite, its ventral margin forming an even curve of ca. 140º, except for the slender apical part without any ventral or lateral processes, ridges, invaginations, etc. (as is generally found in the Eumerini), with a pair of medially directed triangular extensions of the dorsal margins, its dorsal surface otherwise membranous from just posterior to the dorsomedian process for entire length, the proximal half dorsolaterally with a fine granulose sculpture, with fine, oblique transverse striae present laterally and ventrally, but most parts of hypandrium more or less smooth, no trace of a suture discernible where the superior lobes are fused with the hypandrium, the narrow apical part [i.e. caudal to the (apparently bare) lingula region] with a subapical, well-developed ctenidion, apical half deeply invaginated ventrally, its median wall with a densely pilose, triangular ventral process, otherwise simple. Phallus extremely long, fine and hair-like from just posterior to its base to the apex, the two ducti separated a short distance from the phallic base, all the other sclerotized structures present in Eumerini (as in almost all Syrphidae) reduced to a completely unsclerotized, apparently featureless membrane, phallic apodeme small, pump small, very heavily sclerotized. The hamus is entirely absent.
.
Lyneborgimyia magnifica Doczkal and Pape, 2009.
Doczkal, D. and Pape, T. (2009). Lyneborgimyia magnifica gen. et sp.n. (Diptera: Syrphidae) from Tanzania, with a phylogenetic analysis of the Eumerini using new morphological characters. Systematic Entomology 34, 559-573.
unless otherwise stated